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The difference between 5.56x45mm NATO and .223 Rem is so negligible that it's almost entirely academic. The vast majority of rifles that can use one can use the other. You can't buy separate dies for reloading, and they don't have separate sections in reloading manuals. Yes, 5.56 can produce much higher chamber pressures when fired in a .223 chamber, but in most cases you aren't going to have problems.
And as far as AR-15s... They're modular, easy to work on, parts are readily available. That's what makes it the most popular rifle platform in the US, period. The Mini-14, for example, is the IP of Sturm, Ruger & Co; they're the only ones that make the rifle. An AR-15 is an AR-15, almost regardless of who makes it (other than BCA or PSA). That's nearly unique among firearms; there really aren't any other guns on the market where exactly the same firearm, made to the same set of specs, is made by many different manufacturers. Only Glock makes the Glock 17. Only Sig makes the P320 (so far, despite it being the Army service pistol), and only Beretta makes the model 92/M9; that's why you see so, so many different choices in pistols, because there's not any single standard design that's all made to exactly the same spec. (And, BTW, 1911s are awful in that way, as are all AKs; everything needs to be hand fit.)