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If the only way to stop a school shooter were to kill them, I think most people would do so even if they were not personally threatened.
And many people, including myself, think it is moral to kill even an innocent person if necessary to prevent the death of a greater number of people. That's the trolley problem in a nutshell.
But if I'm willing to kill a person in order to prevent them from killing other people, then I should also be willing to kill a fish in order to prevent it from killing other fish.
Finally, the argument for nonhuman sentience does not turn on the presence or absence of neurons. That would just be a cellular version of speciesism, and it inexplicably eliminates the possibility of sentience in extraterrestrials or machines.
The argument in the OP is based on behaviors, like recognizing self vs nonself, avoiding noxious stimuli, creative problem-solving, etc. Plants do many of these things too, just on longer timescales.